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Mammals and birds recorded in and around Manu National Park, Peru,

Joe Tobias and Nat Seddon, August-October 2001

131 Catharine Street, Cambridge CB1 3AP, UK

joetobias22@hotmail.com

Andean Cock of the Rock	White-throated Jacamar	Collared Puffbird

See full versions of Joes photos at www.worldbirder.com

Itinerary

Flight from Cuzco to Boca Manu (BM), boat to Pakitza guard-station (PK): 02/08/01
Boat from Pakitza guard-station to Cocha Cashu Biological Station: 03/08/01
Cocha Cashu Biological Station (CC): 03/08/01 - 09/10/01
Boat to Boca Manu: 09/10/01
Boat from Boca Manu to Pantiacolla lodge: 10/10/01
Pantiacolla lodge (PA): 10/10/01 - 15/10/01
Boat to Manu Road and transport to Cock-of-the-Rock lodge, San Pedro (SP): 15/10/01
Lift to Pillahuarta (PI): 23/10/01
Lift to Cuzco (CU): 24/10/01
Macchu Picchu (MP): 26/10/01
Travel to Ollantaytambo (OL): 27/10/01
Abra Malaga (AM): 28/10/01

Notes on sites

Cocha Cashu Biological Station (c.300 m): we spent several weeks here, focusing on trails near the lake itself, and several times walking out to (and camping at) "Playa Bonita", a stream some 8 km away along a good trail to the uplands. We spent three morning at the Ccolpa stream, also at the end of a different long trail which leaves the main grid, and four mornings across the Manu river in terra firme forest where there is a trail that you need a boat to reach, plus two mornings around nearby Cocha Totora (reached by trail). We also spent one day at the "pantanal", a seasonally flooded grassy area reachable by trail. As most of the time was spent following a group of pale-winged trumpeters for a research project, canopy flocks received relatively little attention. It is impossible to visit Cocha Cashu without research permission, although Cocha Salvador is just a few kilometres downriver and can be visited by tourists. With research permission, charges for staying at Cashu are around US$11 per day, depending whether the cook is there.

Boca Manu (c.300 m): we never did any birding around here, just saw a few things around the town.

Pantiacolla (c.400 m): we divided our time between floodplain/bamboo trails (e.g. Tinamou and Capybara trails), the large loop (Monk Saki trail) that runs through terra firme, and the ridge trail (Mirador trail) where we spent a night camping at the first Mirador (c.900 m). We paid US$15 per night on special rate and camping; not sure how much full board is.

San Pedro (c.1,500 m): not really a settlement on the Manu Road, more the name of a bridge beside which stands the Cock-of-the-Rock lodge. We stayed on a fine sheltered camping platform at this lodge, which is much cheaper than the rooms. A short (c.400 m) loop trail descends to the river from the garden of the lodge; this passes through some good forest. Below this lodge (along the road) forest becomes somewhat patchy, punctuated by areas of secondary growth and clearings, some of which contain interesting patches of bamboo. Above the lodge, there is good forest interrupted by landslides all the way, past the Cloud Forest lodge (c.1,650 m), and up towards Pillahuarta. There are two roadside Cock-of-the-Rock leks, one "owned by the lower lodge and the other "owned" by the upper lodge. The lower lek (c.150 m uphill from Cock-of-the-Rock lodge) seemed superior in terms of viewing conditions, and is the one referred to in the following accounts. There is an interesting Inca trail starting between the two lodges and heading up to the "raptor watchpoint" (really just a major bend in the road c.500 m above Cloud Forest lodge) and passing through some good habitat. Easier to do it backwards from the watchpoint down.

Pillahuarta (c.2,900 m): a couple of huts by the upper Manu Road. Good birding above and below these huts, where it is possible to camp on the floor if you speak to the old madman across the road.

Cuzco (c.2,800 m): we visited the famous ruins overlooking the town on two occasions, catching a taxi up there.

Macchu Picchu (c.2,600 m): we took the morning train from Cuzco and the bus up to the ruins and then walked down through the woods along a steep trail through stunted forest, catching the train back a few hours later.

Ollantaytambo/Abra Malaga (2,000-4,000 m): bus from Cuzco, then jumped on a truck at dawn going up the Abra Malaga road (they leave every day). This dropped us off at the pass (taking two hours to get there) and we walked over to the (highly trashed) Polylepis groves, then down halfway to Ollantaytambo again before finally getting a lift to town.

Highlights

In terms of birds, pride of place goes to the tame group of Pale-winged Trumpeters that we studied at CC for two months. Other highlights were Solitary Eagle, Scarlet-shouldered Parakeet, Pheasant Cuckoo, Rufous-vented Ground-cuckoo, White-throated Jacamar, Collared Puffbird, Curl-crested Araçari, Rufous-headed Woodpecker, Marcapata Spinetail, White-browed Tit-spinetail, Peruvian Recurvebill, Bamboo Antshrike, Ihering's Antwren, Yellow-rumped Antwren (Threatened), Manu Antbird, White-lined Antbird, Goeldi's Antbird, Red-and-white Antpitta, Rufous-fronted Antthrush (Threatened), Black-faced Cotinga, White-cheeked Tody-tyrant, Ash-breasted Tit-tyrant (Threatened), Cinnamon-faced Tyrannulet, Dusky-tailed Flatbill, Golden-collared Tanager, Chestnut-breasted Mountain-finch, Black-billed Seed-Finch, Yellow-shouldered Grosbeak, Selva Cacique (Threatened) and Pale-eyed Blackbird. In terms of mammals, the most impressive were Hoffmann's Two-toed Sloth, Giant Otter and ten species of monkeys (best of which was Emperor Tamarin).

Species Lists

A total of c.32 mammal species and 670 bird species was encountered. None of the sightings below were made in Manu National Park proper; almost all are within the buffer zone or research zone. Sightings from around Cuzco, Macchu Picchu, Ollantaytambo and Abra Malaga are outside the park entirely. The list is heavily weighted towards the tropical zone as only a small proportion of time was spent at high altitudes. Nomenclature and systematics follows Emmons and Feer (1997) for mammals and the existing Manu bird list (on CC website; with amendations) for birds.

Key:

Site codes match those given under Itinirary.

[.] = note or uncertain record
(T) = tape-recorded

Mammals

Common Opossum  Didelphis marsupialis
Seen on 2-3 occasions at night near camp at CC

Southern Tamandua  Tamandua tetradactyla
Seen on three occasions at CC, twice in the grid and once at "Playa Bonita"

Brown-throated Three-toed Sloth  Bradypus tetradactyla
One encountered crossing the stream at "Playa Bonita", CC, where it was helpless and could be approached to 0 m

Hoffmann's Two-toed Sloth  Choloepus hoffmanni
One seen at night at close range near the camp at CC

Pygmy Marmoset  Cebuella pygmaea
One group was seen on 2-3 occasions by its roost tree near the camp at CC

Saddleback Tamarin  Saguinus fuscicollis
Fairly common at CC, particularly in terra firme (T)

Emperor Tamarin  Saguinus imperator
Seen on c.8 occasions at CC, usually in riverine floodplain forest or bamboo-dominated terra firme

Night Monkey Aotus sp. (either Azara's Night Monkey A. azarae, or Black-headed Night Monkey A. nigriceps)
Fairly frequently seen or heard in the grid at CC, on three occasions in daytime when flushed from roosts in vines or bamboo

Dusky Titi Monkey  Callicebus moloch brunneus
Fairly common, especially by voice, at CC (T)

Common Squirrel Monkey  Saimiri sciureus
Very common at CC (T)

Brown Capuchin Monkey  Cebus apella
Common at CC

White-faced Capuchin Monkey  Cebus albifrons
Common at CC

Red Howler Monkey  Alouatta seniculus
Common, especially by voice, and tame at CC; also seen at PA (T)

Common Woolly Monkey  Lagothrix lagothricha
A group was seen from the first Mirador at PA, and another in forest from the Manu Road above SP

White-bellied Spider Monkey  Ateles belzebuth
Very common at CC and heard at PA (T)

Short-eared Dog  Atelocynus microtis
One set of very fresh tracks was seen along the stream at "Playa Bonita", CC

Crab-eating Raccoon  Procyon cancrivorus
Never seen, but tracks were found regularly at CC

Kinkajou  Potus flavus
c.5 seen at night in the grid at CC

Tayra  Eira barbara
Three individuals were watched at close range as they chased each other in the grid at CC

Giant Otter  Pteronura brasiliensis
A group of nine spent a week or two on the lake at CC; also seen at Cocha Totora

Ocelot Leopardus pardalis and Jaguar Panthera onca
Tracks were seen regularly at CC, and Puma Felis concolor tracks were seen on the road above PI, but no cats were seen during the entire time

Brazilian Tapir  Tapirus terrestris
Many tracks seen at CC (where heard occasionally crashing through undergrowth at night), and one set of tracks seen at PA. Never seen.

Collared Peccary  Tayassu tajacu
Small groups of 1-3 were quite commonly encountered in the grid at CC

White-lipped Peccary  Tayassu pecari
Large herds (containing up to 80 individuals) were regularly encountered in the grid at CC

Red Brocket Deer  Mazama americana
Regularly seen in the grid at CC, where tame

Southern Amazon Red Squirrel  Sciurus spadiceus
Common at CC and PA (T)

Bolivian Squirrel  Sciurus ignitus
2-3 seen at CC. One small Sciurus seen in lower-stratum at PA looked more like sanborni, but this identification is left provisional.

Capybara  Hydrochaeris hydrochaeris
A few family groups seen on sand-banks along the Manu river

Paca  Agouti paca
Regularly seen at night at CC

Brown Agouti  Dasyprocta variegata
Fairly common in the grid at CC

Green Acouchy  Myoprocta pratti
Individuals regularly encountered scampering about in the leaf-litter at CC

Vizcacha  Vizcacha vizcacha
A pair at close range on cliffs next to the Polylepis grove at AM

Brazilian Rabbit  Sylvilagus brasiliensis
One seen at night  in the grid at CC

Birds

Grey Tinamou  Tinamus tao
One flushed at close range at CC, where only heard twice

Black Tinamou  Tinamus osgoodi
One (maybe two) heard from the Mirador at dawn, PA (T)

Great Tinamou  Tinamus major
Commonly heard and seen at CC (T)

White-throated Tinamou  Tinamus guttatus
Heard once or twice at CC

Hooded Tinamou  Nothocercus nigrocapillus
1-2 heard around 1,700 m (above SP), but couldn't be enticed into view on steep slopes (T)

Cinereous Tinamou  Crypturellus cinereus
Fairly commonly heard at CC, almost always in terra firme (T)

Little Tinamou  Crypturellus soui
Seen once in várzea and heard occasionally, usually in terra firme at CC (T)

Brown Tinamou  Crypturellus obsoletus
c.8 heard near the first Mirador at PA, many more seen and heard around SP (T)

Undulated Tinamou  Crypturellus undulatus
Very common at CC throughout floodplain and várzea forest; also PA (T)

Black-capped Tinamou  Crypturellus atrocapillus
Fairly common and responsive to playback along the stream at "Playa Bonita", CC. Also heard by the Madre de Dios river at PA (T)

Variegated Tinamou  Crypturellus variegatus
Only heard twice in the uplands near "Playa Bonita", CC

Bartlett's Tinamou  Crypturellus bartletti
Heard commonly and seen regularly in floodplain forest (very rarely in terra firme) at CC (T)

Least Grebe  Tachybaptus dominicus
Small numbers on the lake at CC

Neotropic Cormorant  Phalacrocorax brasilianus
Small numbers regular around the lake at CC; also along the Manu and Madre de Dios rivers

Anhinga  Anhinga anhinga
Small numbers regular at the lake at CC; also along the Manu and Madre de Dios rivers

Horned Screamer  Anhima cornuta
Regular and noisy on sand-banks along the Manu river, and at Cocha Totora, CC (T)

Orinoco Goose  Neochen jubata
Regular on sand-banks along the Manu river

Muscovy Duck  Cairina moschata
Small numbers regular on the lake at CC; also on sand-banks along the Manu river

Torrent Duck  Merganetta armata
Only seen along the river near Macchu Picchu, where this species seemed common

Snowy Egret  Egretta thula
Small numbers along Madre de Dios river

Capped Heron  Pilherodius pileatus
Regular around the lake at CC, and along rivers

Cocoi Heron  Ardea cocoi
Regular around the lake at CC, and fairly common along all rivers

Great Egret  Casmerodius albus
Regular around the lake at CC, and along rivers

Cattle Egret  Bubulcus ibis
One flock seen in flight near Boca Manu

Striated Heron  Butorides striatus
Fairly common around the lake at CC, and along rivers

Agami Heron  Agamia agami
Seen c.15 times around the lake at CC, and along small streams within forest nearby

Boat-billed Heron  Cochlearius cochlearius
Small numbers seen at night around the lake at CC, and along the stream at "Playa Bonita"

Fasciated Tiger-Heron  Tigrisoma fasciatum
c.10 seen while travelling along the Madre de Dios river in the PA area

Rufescent Tiger-Heron  Tigrisoma lineatum
Fairly common around the lake at CC

Green Ibis  Mesembrinibis cayennensis
Small numbers regular around the lake at CC

Roseate Spoonbill  Ajaia ajaja
One flock of c.15 on a sand-bank downstream of PA on the Madre de Dios river

Wood Stork  Mycteria americana
Small groups seen occasionally along the Manu river

Black Vulture  Coragyps atratus
Small numbers seen throughout

Turkey Vulture  Cathartes aura
Recorded from the lower Manu Road, up to above SP and in the CU area

Greater Yellow-headed Vulture  Cathartes melambrotus
Fairly common around CC

King Vulture  Sarcoramphus papa
Small numbers recorded at CC, mostly at "Playa Bonita"

Osprey  Pandion haliaetus
Occasional at the lake at CC and along major rivers to PA

Grey-headed Kite  Leptodon cayanensis
One watched gliding over the Manu River at CC

Swallow-tailed Kite  Elanoides forficatus
Regular in small numbers at CC, PA and SP

Snail Kite  Rostrhamus sociabilis
Small numbers occasional at CC and nearby Cocha Totora

Double-toothed Kite  Harpagus bidentatus
Regularly seen perched or circling at CC and PA; also one at c.1200 m, below SP

Plumbeous Kite Ictinia plumbea
Regular at CC, PA and below SP

Tiny Hawk  Accipiter superciliosus
One watched flying across the Manu river at CC

Plain-breasted Hawk  Accipiter ventralis
One seen above PI

Slate-colored Hawk  Leucopternis schistacea
Fairly common (at least by voice) in várzea, floodplain and swampy areas near CC (T)

White-browed Hawk  Leucopternis kuhli
Seen twice in terra firme across the Manu river from CC, and once near "Playa Bonita"

Great Black-Hawk  Buteogallus urubitinga
Small numbers seen along the Manu river

Black-collared Hawk  Busarellus nigricollis
Fairly common around CC and nearby Totora

Solitary Eagle  Harpyhaliaetus solitarius
One seen circling over hillside opposite SP

Roadside Hawk  Buteo magnirostris
Fairly common throughout lowlands and up to SP (T)

Short-tailed Hawk  Buteo brachyurus
One seen flying over the confluence of the Manu and Madre de Dios rivers at BM

White-throated Hawk  Buteo albigula
At least two above PI

Variable Hawk  Buteo polyosoma
A few around AM

Black-and-white Hawk-Eagle  Spizastur melanoleucus
One seen soaring over forested hillsides at c.1,300 m on the Manu Road

Ornate Hawk-Eagle  Spizaetus ornatus
Fairly common at CC where two active nests were found (T)

Black Caracara  Daptrius ater
Fairly common at CC and PA (T)

Red-throated Caracara  Daptrius americanus
Common and very noisy around CC and PA (T)

Laughing Falcon  Herpetotheres cachinnans
Regularly heard at dawn and dusk at CC, but never seen (T)

Barred Forest-Falcon  Micrastur ruficollis
Regularly heard at CC and seen on c.5 occasions; also heard near the first Mirador at PA (T)

Lined Forest-Falcon  Micrastur gilvicollis
Heard on c.3 occasions around CC, and played back into view once (T)

Collared Forest-Falcon  Micrastur semitorquatus
Seen on c.5 occasions around CC (possible that one of these was buckleyi, but views were inadequate)

American Kestrel  Falco sparverius
Seen above PI and around CU

Bat Falcon  Falco rufigularis
Fairly common around the lakes and rivers at CC (T)

Speckled Chachalaca  Ortalis guttata
Regular in lakeside or riverside growth at CC and PA (T)

Andean Guan  Penelope montagnii
Small numbers seen above SP

Spix's Guan  Penelope jacquacu
Fairly common around CC (T)

Blue-throated Piping-Guan  Pipile cumanensis
Fairly common around CC and PA (T)

Razor-billed Curassow  Mitu tuberosa
Fairly common around CC: seen or heard most days (T)

Marbled Wood-Quail  Odontophorus gujanensis
Heard on 2-3 occasions in riverine growth along the Manu river, but never seen

Starred Wood-Quail  Odontophorus stellatus
Heard regularly around the grid at CC, and on the path to the uplands, and seen on c.5 occasions (T)

Rufous-sided Crake  Laterallus melanophaius
Fairly common around the lakes at Cashu and Totora, where vocal and easily seen at dawn or dusk coming out onto floating vegetation (T)

Grey-breasted Crake  Laterallus exilis
Heard infrequently at CC and Totora and never seen

Grey-necked Wood-Rail  Aramides cajanea
Fairly common at the fringes of CC (T)

Blackish Rail  Pardirallus nigricans
One from a bus near OL

Purple Gallinule  Porphyrio martinicus
A feww around CC

Common Moorhen  Gallinula chloropus
A few around CC

Sungrebe  Heliornis fulica
Regular around the shores of CC and once along the stream at "Playa Bonita"

Sunbittern  Eurypyga helias
Regular around the lake at CC, along the stream at "Playa Bonita" and one seen on the Manu Road where a bridge crosses a river at c.1,250 m (T)

Limpkin  Aramus guarauna
Regular around the lake at CC

Pale-winged Trumpeter  Psophia leucoptera
Regularly seen around CC, where there is one very tame group, and alarm calls heard once at PA (T)

Wattled Jacana  Jacana jacana
Fairly common at CC, common at Cocha Totora (T)

Greater Yellowlegs  Tringa melanoleuca
Two seen at the mouth of the Ccolpa stream, and small numbers elsewhere along the Manu and Madre de Dios rivers

Lesser Yellowlegs  Tringa flavipes
A few (c.5) seen at the mouth of the Ccolpa stream, CC

Solitary Sandpiper  Tringa solitaria
Small numbers along the Manu river, and the Ccolpa and "Playa Bonita" streams, CC

Spotted Sandpiper  Tringa macularia
Fairly common along major rivers from CC to PA

Pectoral Sandpiper  Calidris melanotos
Small numbers seen at the mouth of the Ccolpa stream and on sand-banks along the Manu river

Buff-breasted Sandpiper  Tryngites subruficollis
One seen on a grassy sandbank along the Manu river

White-backed Stilt  Himantopus melanurus
One small flock downstream of PA on the Madre de Dios river

Collared Plover  Charadrius collaris
Fairly common on sand-banks along the Manu river

Pied Lapwing  Vanellus cayanus
Fairly common along the Manu river

Andean Lapwing  Vanellus resplendens
2-3 in the open valley floor at AM

Andean Gull  Larus serranus
One seen flying over fields near the pass at AM

Large-billed Phaetusa simplex
Large numbers breeding on sand-banks along the Manu river

Yellow-billed Tern  Sterna superciliaris
Small numbers breeding on sand-banks along the Manu river

Black Skimmer  Rynchops niger
Several fairly large colonies along the Manu river

Spot-winged Pigeon  Columba maculosa
A few around CU

Band-tailed Pigeon  Columba fasciata
Regular in cloud forest above SP and around PI

Pale-vented Pigeon  Columba cayennensis
Common in lakeside or riverside forest at CC and PA (T)

Plumbeous Pigeon  Columba plumbea
Common at CC, PA and right up to nearly 2,000 m on Manu Road (T)

Ruddy Pigeon  Columba subvinacea
Common at CC and PA (T)

Eared Dove  Zenaida auriculata
Common around CU and OL

Blue Ground-Dove  Claravis pretiosa
Occasional at PK and CC (T)

Grey-fronted Dove  Leptotila rufaxilla
Common, particularly in waterside forest, at CC and PA (T)

Sapphire Quail-Dove  Geotrygon saphirina
One seen walking on forest floor a short way up the Mirador trail at PA

White-throated Quail-Dove  Geotrygon frenata
Several calling around PI, others heard at SP were probably this species (T)

Ruddy Quail-Dove  Geotrygon montana
Fairly common at CC and PA (T)

Blue-and-yellow Macaw  Ara ararauna
Common at CC and PA (T)

Scarlet Macaw  Ara macao
Regular at CC, several pairs nesting in palms in swampy areas (T)

Red-and-green Macaw  Ara chloropterus
Regular at CC and PA

Chestnut-fronted Macaw  Ara severa
Fairly common at CC and PA (T)

Blue-headed Macaw  Ara couloni
Never seen well, but it's double note was heard on a few occasions over forest at CC

White-eyed Parakeet  Aratinga leucophthalmus
A few at CC and PA

Dusky-headed Parakeet  Aratinga weddellii
Fairly common at CC and PA (T)

Mitred Parakeet  Aratinga mitrata
Only identified below Macchu Picchu where it seemed common

Painted Parakeet  Pyrrhura picta
Common at CC and PA; never saw rupicola and this was definitely in the minority (T)

Andean Parakeet  Bolborhynchus orbygnesius
One flock of c.25 at AM

Dusky-billed Parrotlet  Forpus sclateri
Fairly common along the streams near the Ccolpa and at "Playa Bonita" where they visit clay-licks; never seen in the forest itself (T)

Cobalt-winged Parakeet  Brotogeris cyanoptera
Very common throughout the forest at CC and PA (T)

Tui Parakeet  Brotogeris sanctithomae
Regular around the lakeshore and riverside at CC (T)

Scarlet-shouldered Parrotlet  Touit huetii
One pair seen at PA where digging a cavity into an arboreal termitary (T)

White-bellied Parrot  Pionites leucogaster
Common and vocal at CC and PA (T)

Blue-headed Parrot  Pionus menstruus
Fairly common and vocal at CC and PA (T)

Yellow-crowned Parrot  Amazona ochrocephala
Regular around CC (T)

Scaly-naped Parrot  Amazona mercenaria
Flocks flying at dusk over SP daily

Mealy Parrot  Amazona farinosa
Common and noisy at CC and PA (T)

Yellow-billed Cuckoo  Coccyzus americanus
Two seen with canopy flocks towards the end of our stay at CC

Dark-billed Cuckoo  Coccyzus melacoryphus
One seen in lakeside trees at CC, another seen in floodplain forest at PA

Squirrel Cuckoo  Piaya cayana
Common in lower-lying areas (T)

Hoatzin  Opisthocomus hoazin
Common around CC and Totora (T)

Greater Ani  Crotophaga major
Fairly common around CC and Totora

Smooth-billed Ani  Crotophaga ani
Seen around PA and the lower Manu Road

Pheasant Cuckoo  Dromococcyx phasianellus
Regularly heard in bamboo along the stream at "Playa Bonita", where seen once very well in response to playback (T)

Pavonine Cuckoo  Dromococcyx pavoninus
Heard twice in bamboo in terra firme across the river from CC (T)

Rufous-vented Ground-Cuckoo  Neomorphus geoffroyi
Prolonged views at close range by the station at CC, and at the beginning of the Tinamou trail at PA

Barn Owl  Tyto alba
One seen in flight shortly after dusk at PI

Tropical Screech-Owl  Otus choliba
Heard twice at the lake at CC

Rufescent Screech-Owl  Otus ingens
Heard regularly around SP and responded strongly to playback in the grounds of the lodge (T)

Tawny-bellied Screech-Owl  Otus watsonii
Heard commonly at CC and PA; seen once each at both these sites (T)

Vermiculated Screech-Owl  Otus vermiculatus
2-3 heard near the top of the Mirador trail, PA (T)

Mottled Owl  Strix virgata
Heard occasionally in forest at CC and once at PA (T)

Black-banded Owl  Strix huhula
Heard once at CC

Crested Owl  Lophostrix cristata
Heard nightly at CC and PA; seen three times at close range in daylight at CC (T)

Spectacled Owl  Pulsatrix perspicillata
Heard often at CC, and seen once in daylight (T)

Band-bellied Owl  Pulsatrix melanota
Two pairs heard near the first Mirador at PA; came very close to playback but couldn't be seen (T)

Amazonian Pygmy-Owl  Glaucidium hardyi
Commonly heard and regularly seen in daylight at CC (T)

Ferruginous Pygmy-Owl  Glaucidium brasilianum
Heard occasionally at CC, mostly at "Playa Bonita" (T)

Great Potoo  Nyctibius grandis
Heard occasionally at CC but never seen (T)

Long-tailed Potoo  Nyctibius aethereus
Heard on c.5 occasions at CC but never seen; one found incubating eggs at PA (T)

Grey Potoo  Nyctibius griseus
Heard twice at CC in riverine Cecropias

Short-tailed Nighthawk Lurocalis semitorquatus
Seen at dusk from a canoe flying over forest at CC

Rufous-bellied Nighthawk  Lurocalis rufiventris
One bird played in to close range at dusk at PI (T)

Sand-colored Nighthawk  Chordeiles rupestris
Common on sand-banks along the Manu river, and several visited the lake itself at CC each dusk

Pauraque  Nyctidromus albicollis
Fairly common around the lake at CC, and even quite deep in the forest (T0

Ocellated Poorwill  Nyctiphrynus ocellatus
Common in forest at CC: often heard and c.10 nests found by eyeshine at night (T)

Lyre-tailed Nightjar  Uropsalis lyra
2-3 heard at c.1,700 m above SP; two males seen well and one female found in daylight on a roadside nest (T)

Chestnut-collared Swift  Streptoprocne rutila
Common in large flocks at SP

White-collared Swift  Streptoprocne zonaris
Occasional over CC and fairly common along the Manu Road

Grey-rumped Swift  Chaetura cinereiventris
Fairly common at CC

Pale-rumped Swift  Chaetura egregia
Regular at CC, in small numbers over the "Playa Bonita" stream, and seen once at PA

Short-tailed Swift  Chaetura brachyura
Fairly common at CC

Fork-tailed Palm-Swift  Tachornis squamata
Small numbers seen at CC

Rufous-breasted Hermit  Glaucis hirsuta
Common at CC where several nests were found

Pale-tailed Barbthroat  Threnetes leucurus
A few seen at CC and several seen at PA (T)

Green Hermit  Phaethornis guy
2-3 seen at SP

Great-billed Hermit  Phaethornis malaris
Seen rarely, or at least identified rarely in terra firme at CC

White-bearded Hermit  Phaethornis hispidus
Common and vocal at CC (T)

Koepcke's Hermit  Phaethornis koepckeae
Several seen along the Mirador trail at PA

Needle-billed Hermit  Phaethornis philippii
One seen in bamboo near the river on the Tinamou trail at PA

Reddish Hermit  Phaethornis ruber
Common at CC, PA and seen below SP (T)

[White-browed Hermit  Phaethornis stuarti]
Possibly seen on several occasions but we found this species impossible to identify with certainty

Buff-tailed Sicklebill  Eutoxeres condamini
Seen once below SP

Green-fronted Lancebill  Doryfera ludovicae
Seen regularly at flowering trees near SP

Grey-breasted Sabrewing  Campylopterus largipennis
Seen on 2-3 occasions at CC

White-necked Jacobin  Florisuga mellivora
Seen occasionally at CC (mostly feeding high over the lake or streams), also in the clearing at PA

Green Violet-ear  Colibri thalassinus
Small numbers seen between SP and PI

Sparkling Violet-ear  Colibri coruscans
Fairly common between SP and PI (T)

Blue-tailed Emerald  Chlorostilbon mellisugus
A female seen well at CC in August

Fork-tailed Woodnymph  Thalurania furcata
Fairly common at CC and PA

White-chinned Sapphire  Hylocharis cyanus
Fairly common in more open forest or clearings at CC (T)

Golden-tailed Sapphire  Chrysuronia oenone
Seen once at SP

Many-spotted Hummingbird  Taphrospilus hypostictus
Seen once at SP in roadside low canopies, fighting gamely with a Sparkling Violetear

Green-and-white Hummingbird  Amazilia viridicauda
Singing around the ruins at MP

Sapphire-spangled Emerald  Amazilia lactea
A few seen and heard along the lower Manu Road (T)

Speckled Hummingbird  Adelomyia melanogenys
Common and vocal in forest around SP (T)

Gould's Jewelfront  Heliodoxa aurescens
Seen on c.5 occasions at CC

Violet-fronted Brilliant  Heliodoxa leadbeateri
Fairly common at SP, visiting feeders at Cock-of-the-rock lodge

Giant Hummingbird  Patagona gigas
1-2 seen at AM

Shining Sunbeam  Aglaeactis cupripennis
Fairly common at AM

White-tufted Sunbeam  Aglaeactis castelnaudi
c.4 seen in shrubbery by the road between OL and AM

Great Sapphirewing  Pterophanes cyanopterus
c.15 seen below AM

Violet-throated Starfrontlet  Coeligena violifer
1 seen at c.2,000 m (above SP), and another 3-4 seen at c.3,100 m (above PI)

Sword-billed Hummingbird  Ensifera ensifera
One seen well above PI

Chestnut-breasted Coronet  Boissonneaua matthewsii
One seen in woodland below MP

Amethyst-throated Sunangel  Heliangelus amethysticollis
Fairly common in woodland near PI

Coppery-naped Puffleg  Eriocnemis sapphiropygia
One seen at c.2,000 m (above SP)

Booted Racquet-tail  Ocreatus underwoodii
Small numbers seen in roadside vegetation above SP

Green-tailed Trainbearer  Lesbia nuna
A few seen near CU

Scaled Metaltail  Metallura aeneocauda
One seen well at PI

Tyrian Metaltail  Metallura tyrianthina
Race with blue (not reddish) tail common above c.1,800 m on Manu Road

Blue-mantled Thornbill  Chalcostigma stanleyi
c.5 seen around Polylepis at AM

Long-tailed Sylph  Aglaiocercus kingi
Fairly common above SP

Wedge-billed Hummingbird  Schistes geoffroyi
One seen in song c.300 m downhill from Cloud Forest lodge, Manu Road (T)

Black-eared Fairy  Heliothryx aurita
2-3 seen in canopies at CC

Long-billed Starthroat  Heliomaster longirostris
One at CC (edge of Cocha Totora)

White-bellied Woodstar  Acestrura mulsant
One male at SP

Crested Quetzal  Pharomachrus antisianus
2-3 between SP and Cloud Forest lodge (T)

Golden-headed Quetzal  Pharomachrus auriceps
c.8 seen above SP and below PI; quite often heard (T)

Black-tailed Trogon  Trogon melanurus
Very common at CC, less so at PA (T)

White-tailed Trogon  Trogon viridis
Scarce at CC, more common at PA (T)

Collared Trogon  Trogon collaris
Fairly common at CC and PA

Masked Trogon  Trogon personatus
One in song above PI (T)

Blue-crowned Trogon  Trogon curucui
Fairly common at CC and a few at PA (T)

Violaceous Trogon  Trogon violaceus
Scarce at CC, more common at PA and lower Manu Road (T)

Ringed Kingfisher  Ceryle torquata
Common at lakes and rivers in the lowlands

Amazon Kingfisher  Chloroceryle amazona
A few around CC and along the Manu and Madre de Dios rivers

Green Kingfisher  Chloroceryle americana
Fairly common at CC, including along small forest streams

Green-and-rufous Kingfisher  Chloroceryle inda
Seen occasionally at CC

American Pygmy Kingfisher  Chloroceryle aenea
Seen occasionally at CC

Broad-billed Motmot  Electron platyrhynchum
Seen c.5 times and heard fairly often at CC (T)

[Rufous Motmot Baryphthengus martii]
Probably heard at PA, but never definitely recorded

Blue-crowned Motmot  Momotus momota
Common at CC and PA (T)

Highland Motmot  Momotus aequatorialis
Seen on c.4 occasions at SP

Chestnut Jacamar  Galbalcyrhynchus purusianus
Heard regularly and seen on c.6 occasions in swampy or lakeside forest at CC

White-throated Jacamar  Brachygalba albogularis
Fairly common along the Ccolpa and "Playa Bonita" streams, but not seen anywhere else at CC (T)

Bluish-fronted Jacamar  Galbula cyanescens
Common, especially in streamside, riverside or lakeside vegetation at CC; seen up to around 1,400 m on the Manu road (T)

Great Jacamar  Jacamerops aureus
Heard regularly and seen occasionally at CC (T)

White-necked Puffbird  Notharchus macrorhynchos
Never seen, but occasionally heard calling from upper canopy at CC (T)

Chestnut-capped Puffbird  Bucco macrodactylus
Only seen once, at a clearing in the grid at CC

Collared Puffbird  Bucco capensis
Seen on c.5 occasions in the grid at CC, usually at close range in lower-stratum; heard only once, after dusk (T)

Striolated Puffbird  Nystalus striolatus
Heard regularly at CC and seen on c.5 occasions, usually in mid-levels of vine-tangled forest near the Manu river

Semicollared Puffbird  Malacoptila semicincta
Seen on c.8 occasions in lower-stratum at CC

Rufous-capped Nunlet  Nonnula ruficapilla
Seen regularly in riverine regrowth along the Manu river and smaller streams (e.g. at "Playa Bonita" and Ccolpa)

Black-fronted Nunbird  Monasa nigrifrons
Very common in floodplain forest at CC and PA (T)

White-fronted Nunbird  Monasa morphoeus
Scarce in floodplain forest and slightly more common in terra firme at CC and PA (T)

Swallow-winged Puffbird  Chelidoptera tenebrosa
Common in riverside vegetation throughout the lowlands; many nests found along the "Playa Bonita" and Ccolpa streams (T)

Gilded Barbet  Capito auratus
Very common by voice in canopies at CC and PA, much less often seen (T)

Lemon-throated Barbet  Eubucco richardsoni
Fairly common by voice at CC and PA, seen regularly

Scarlet-hooded Barbet  Eubucco tucinkae
1-2 seen along the "Playa Bonita" stream, and several in riverside Cecropias at PA

Versicolored Barbet  Eubucco versicolor
2-3 seen above SP, around 2,200 m

Emerald Toucanet  Aulacorhynchus prasinus
A few at CC, mostly around the "Playa Bonita" and Ccolpa streams

Chestnut-tipped Toucanet  Aulacorhynchus derbianus
Not seen, but one taped while calling on the Manu Road at c.1,200 m

Blue-banded Toucanet  Aulacorhynchus coeruleicinctis
c.8 seen above SP, between 1,750 and 2,400 m

Lettered Aracari  Pteroglossus inscriptus
Fairly common at CC and PA

Brown-mandibled Aracari  Pteroglossus mariae
Fairly common at CC

Chestnut-eared Aracari  Pteroglossus castanotis
Common at CC and PA (T)

Curl-crested Araçari  Pteroglossus beauharnaesii
Seen on c.15 occasions at CC

Golden-collared Toucanet  Selenidera reinwardtii
Fairly common by voice at CC, but only seen on c.10 occasions

Channel-billed Toucan  Ramphastos vitellinus
Fairly common by voice at CC (T)

White-throated Toucan  Ramphastos tucanus
Very common at CC and fairly common at PA (T)

Rufous-breasted Piculet  Picumnus rufiventris
Seen or heard on c.10 occasions in swampy or bamboo-dominated forest at CC (T)

Yellow-tufted Woodpecker  Melanerpes cruentatus
Common at CC and PA (T)

Little Woodpecker  Veniliornis passerinus
Fairly common at CC, PA, and the lower Manu Road, mostly in riverside habitat (T)

Red-stained Woodpecker  Veniliornis affinis
Fairly common with mid-stratum flocks at CC

[White-throated Woodpecker Piculus leucolaemus]
Possibly seen at CC, but all good views were of the next species

Golden-green Woodpecker  Piculus chrysochloros
Often seen and heard at CC (T)

Golden-olive Woodpecker  Piculus rubiginosus
Seen on c.8 occasions above SP

Spot-breasted Woodpecker  Colaptes punctigula
Seen or heard on c.4 occasions in riverine or lakeside habitat at CC (T)

Andean Flicker  Colaptes rupicola
Fairly common at AM

Scaly-breasted Woodpecker  Celeus grammicus
Regularly seen, and heard almost daily at CC (T)

Chestnut Woodpecker  Celeus elegans
Seen on c.15 occasions at CC, and once at PA (T)

Cream-colored Woodpecker  Celeus flavus
Seen or heard on c.10 occasions at CC, always around swampy areas of floodplain forest in grid (T)

Rufous-headed Woodpecker  Celeus spectabilis
Heard on most visits to the "Playa Bonita" stream, where it was seen on 3 occasions in streamside cane and bamboo (T)

Lineated Woodpecker  Dryocopus lineatus
Occasionally heard in flooded areas with dead trees at CC, also at PA and lower Manu Road

Red-necked Woodpecker  Campephilus rubricollis
Fairly common at CC and PA (T)

Crimson-crested Woodpecker  Campephilus melanoleucos
Fairly common - but less so than the last species - at CC

Plain-brown Woodcreeper  Dendrocincla fuliginosa
Fairly common, at least by voice, at CC and PA (T)

White-chinned Woodcreeper  Dendrocincla merula
Seen or heard (a quiet tittering) on c.20 occasions at CC - accompanying most ant-swarms (T)

Long-tailed Woodcreeper  Deconychura longicauda
1-2 heard most days in the grid at CC (T)

Olivaceous Woodcreeper  Sittasomus griseicapillus
Regularly seen and heard accompanying mid-stratum mixed-species flocks at CC and PA (T)

Wedge-billed Woodcreeper  Glyphorynchus spirurus
Fairly common at CC, but infrequently heard

Long-billed Woodcreeper  Nasica longirostris
Regular around the lake at CC (T)

Cinnamon-throated Woodcreeper  Dendrexetastes rufigula
Regularly heard at CC and PA, but only seen on c.5 occasions at the former (T)

Strong-billed Woodcreeper  Xiphocolaptes promeropirhynchus
Lawrence's Thrushes Turdus lawrencii were regularly heard imitating this species at CC, but the real thing was only heard twice

Barred Woodcreeper  Dendrocolaptes certhia
Fairly common at CC, especially by voice (T)

Black-banded Woodcreeper  Dendrocolaptes picumnus
Appeared much less common than the previous species, only seen or heard on c.3 occasions at CC

Straight-billed Woodcreeper  Xiphorhynchus picus
Common in lakeside or riverside habitat at CC and PA (T)

Striped Woodcreeper  Xiphorhynchus obsoletus
Heard regularly in swampy woodland with stands of Heliconia, or in lakeside vegetation at CC (T)

Tschudi's Woodcreeper  Xiphorhynchus chunchotambo
Definitely saw this twice at CC, and it was probably heard occasionally

Elegant Woodcreeper  Xiphorhynchus elegans
Commonly heard and regularly seen with lower-stratum flocks at CC and PA (T)

Buff-throated Woodcreeper  Xiphorhynchus guttatus
Very common and vocal at CC and PA (T)

Olive-backed Woodcreeper  Xiphorhynchus triangularis
Seen on c.3 occasions with mixed-species flocks above SP

Lineated Woodcreeper  Lepidocolaptes albolineatus
Seen regularly accompanying canopy flocks at CC and PA, heard infrequently (T)

Red-billed Scythebill  Campylorhamphus trochilirostris
Seen in bamboo-dominated streamside woodland at "Playa Bonita" and in taller terra firme at PA (T)

Bar-winged Cinclodes  Cinclodes fuscus
Fairly common at AM

White-winged Cinclodes  Cinclodes atacamensis
2-3 at AM

Pale-legged Hornero  Furnarius leucopus
Fairly common at CC and PA in lakeside, riverside or streamside habitats (T)

Azara's Spinetail  Synallaxis azarae
Fairly common in roadside growth at SP and towards PI (T)

Plain-crowned Spinetail  Synallaxis gujanensis
A few seen and heard in tangled edge habitat at the "pantanal" and "Playa Bonita" at CC (T)

Rufous Spinetail  Synallaxis unirufa
2-3 seen above SP

Ash-browed Spinetail  Cranioleuca curtata
Fairly common with mixed-species flocks at and above SP, also seen on two occasions high on the Mirador trail at PA (T)

Marcapata Spinetail  Cranioleuca marcapatae
2-3 active nests found on the long flat stretch of road a few kms above PI; this species seems to favour building untidy nests dangling over openings such as this road (T)

Speckled Spinetail  Cranioleuca gutturata
2-3 seen at CC in tangled streamside trees; a few also seen lower down at PA

White-browed Tit-spinetail  Leptasthenura xenothorax
c.4 seen in Polylepis at AM

Tawny Tit-spinetail  Leptasthenura yanacensis
Fairly in and around Polylepis groves at AM

Streak-throated Canastero  Asthenes humilis
Fairly common at AM

Line-fronted Canastero  Asthenes urubambensis
2-3 seen at AM

Plain Softtail  Phacellodomus fusciceps
Only one seen at CC in lakeside tangled vines 

Spotted Barbtail  Premnoplex brunnescens
Fairly common by voice at SP and seen accompanying low flocks on c.6 occasions (T)

Pearled Treerunner  Margarornis squamiger
A few seen with mixed-species flocks at PI

Sharp-tailed Streamcreeper  Lochmias nematura
One bird heard and seen near a stream on the Mirador trail at PA

Streaked Tuftedcheek  Pseudocolaptes boissonneautii
A few seen at PI

Chestnut-winged Hookbill  Ancistrops strigilatus
Seen accompanying mixed-species flocks at CC on c.8 occasions

Striped Woodhaunter  Hyloctistes subulatus
Fairly common by voice at CC, and seen on c.10 occasions, sometimes with flocks but often independent (T)

Buff-browed Foliage-gleaner  Syndactyla rufosuperciliata
2-3 seen and heard above SP, around 1,750 m (T)

Montane Foliage-gleaner  Anabacerthia striaticollis
Fairly common along the Manu Road, around and above SP, usually with every mixed-species flock (T)

Rufous-tailed Foliage-gleaner  Philydor ruficaudatus
Fairly common with mid-stratum foraging flocks at CC (T)

Rufous-rumped Foliage-gleaner  Philydor (erythrocercus) lyra
One seen with a mixed-species flock at c.1,600 m near SP

Rufous-rumped Foliage-gleaner  Philydor erythrocercus ochrogaster
Fairly common with mid-stratum flocks at CC and PA (T)

Chestnut-winged Foliage-gleaner  Philydor erythrpterus
Seen on c.5 occasions with mid-stratum foraging flocks at CC

Buff-fronted Foliage-gleaner  Philydor rufus
2-3 seen or heard in Cecropia trees and Gynerium cane along the Manu river

Peruvian Recurvebill  Simoxenops ucayalae
Fairly common (at least by voice) in bamboo at "Playa Bonita" and in the nearby uplands at CC (T)

Striped Treehunter  Thripadectes holostictus
Fairly common around and below SP in lower growth and mixed-species flocks (T)

Buff-throated Foliage-gleaner  Automolus ochrolaemus
Heard regularly (and occasionally seen) at CC, almost always in bamboo-dominated or terra firme forest (T)

Bamboo Foliage-gleaner  Automolus dorsalis
Regularly heard and seen along the "Playa Bonita" stream at CC, and appeared common in bamboo at PA (T)

Olive-backed Foliage-gleaner  Automolus infuscatus
Fairly common by voice in floodplain forest at CC and PA, where sometimes encountered with lower-stratum flocks (T)

Brown-rumped Foliage-gleaner  Automolus melanopezus
Scarce in bamboo-dominated terra firme forest at CC (once across the Manu river, and several times in the uplands near "Playa Bonita"; easily picked up by song (T)

Chestnut-crowned Foliage-gleaner  Automolus rufipileatus
Fairly common (at least by voice) in bamboo and cane-dominated regrowth along the "Playa Bonita" stream (T)

Tawny-throated Leaftosser  Sclerurus mexicanus
Heard twice - and seen once after playback - in the grid at CC (T)

Grey-throated Leaftosser  Sclerurus albigularis
One seen and heard just outside the grid at CC on the path to the Ccolpa (T)

Black-tailed Leaftosser  Sclerurus caudacutus
Seen twice and heard on c.5 occasions at dawn or dusk in the grid at CC (T)

[Rufous-tailed Xenops  Xenops milleri]
At least one bird seen with a canopy flock was almost certainly this species, but unfortunately the views were not good enough to be 100% sure

Plain Xenops  Xenops minutus
Common at CC and PA, where regularly seen with mixed-species foraging flocks from the undergrowth to subcanopy (T)

[Slender-billed Xenops  Xenops tenuirostris]
Severa l proables seen in the canopy accompanying mixed-species flocks, but views were always inconclusive

Streaked Xenops  Xenops rutilans
Fairly common around SP in mixed-species flocks (T)

Fasciated Antshrike  Cymbilaimus lineatus
Fairly common (especially by voice) accompanying mixed-species foraging flocks at CC and PA (T)

Bamboo Antshrike  Cymbilaimus sanctaemariae
Fairly common in bamboo near "Playa Bonita" at CC, and near the Madre de Dios river at PA. Also heard once in bamboo at c.1,000 m from the Manu Road below SP (T)

Great Antshrike  Taraba major
Fairly common in riverine or lakeside regrowth at CC and PA (T)

Chestnut-backed Antshrike  Thamnophilus palliatus
Regularly heard and occasionally seen in roadside shrubbery near SP (T)

White-shouldered Antshrike  Thamnophilus aethiops
Fairly common by voice at CC, and one pair seen at PA (T)

Upland Antshrike  Thamnophilus aroyae
2-4 seen or heard in roadside shrubbery above SP (T)

Plain-winged Antshrike  Thamnophilus schistaceus
Common and vocal at CC and PA where found accompanying almost all mixed-species flocks in the mid-stratum or sub-canopy; we never found Mouse-coloured Antshrike Thamnophilus murinus, and this species must be quite rare in the area (T)

Spot-winged Antshrike  Pygiptila stellaris
Commonly heard and regularly seen at CC and PA, accompanying mixed-species flocks in mid-stratum and canopy (T)

Russet Antshrike  Thamnistes anabatinus
One seen with a mixed-species flock midway up the ridge on the Mirador trail, PA

Plain Antvireo  Dysithamnus mentalis
Never at CC, but a few at PA (T)

Dusky-throated Antshrike  Thamnomanes ardesiacus
Accompanying all lower-stratum flocks at CC and PA (T)

Bluish-slate Antshrike  Thamnomanes schistogynus
Like the previous species, a noisy and conspicuous component of all lower-stratum flocks at CC (T)

Pygmy Antwren  Myrmotherula brachyura
Common at CC and PA, mostly in more vine-tangled situations where it accompanies mixed-species flocks at mid-levels (T)

Sclater's Antwren  Myrmotherula sclateri
Regularly heard at CC, rarely seen, always with canopy flocks, regularly together with Pygmy Antwren; sclateri comes lower in viny areas, where it is occasionally seen alongside its congener - most sclateri didn't have streaky chests, but were easily identified by unique voice and lack of white throat (T)

Amazonian Streaked Antwren  Myrmotherula multostriata
Fairly common (at least by voice) in lakeside vegetation at CC (T)

Stripe-chested Antwren  Myrmotherula longicauda
Regularly heard and occasionally seen around  the lodge at SP (T)

Plain-throated Antwren  Myrmotherula hauxwelli
Regularly seen and heard close to the ground at CC and PA, regularly with lower-stratum flocks (T)

White-eyed Antwren  Myrmotherula leucophthalma
Fairly common in lower-stratum mixed-species flocks; seen every two or three days at CC, also seen at PA (T)

Ornate Antwren  Myrmotherula ornata
Always seen in association with bamboo in this region, and best picked up by voice. Regularly found in or near bamboo patches near "Playa Bonita", CC, and in the bamboo patches downhill from SP, at c.1,300 m (T)

Rufous-tailed Antwren  Myrmotherula erythrura
Only seen at PA, where seemed quite common in mixed-species flocks in lower and mid-strata (T)

White-flanked Antwren  Myrmotherula axillaris
Fairly common component of lower-stratum flocks at CC and PA (T)

Long-winged Antwren  Myrmotherula longipennis
Fairly common in lower-stratum flocks (one pair seemed to accompany most of them) at CC (T)

Ihering's Antwren  Myrmotherula iheringi
Confirmed sightings on four occasions at CC (once in the grid, twice in terra firme across the Manu river and once towards the uplands), and once at PA not far from the lodge on Monk Saki trail; always with small groups of insectivores in lower or mid-strata and best picked up by voice (T)

Grey Antwren  Myrmotherula menetriesii
Fairly common member of lower and mid-strata flocks at CC and PA; seemed commonest in riverine of lakeside forest (T)

Banded Antwren  Dichrozona cincta
Scarce at CC, where seen on c.5 occasions (only once without playback), most regularly along the long trail to Playa Bonita" (T)

Rufous-winged Antwren  Herpsilochmus rufimarginatus
Regularly heard in canopy flocks at CC, but rarely seen (T)

Yellow-breasted Antwren  Herpsilochmus axillaris
One pair was seen on several occasions with a canopy flock just uphill of the Cock-of-the-Rock lek at SP (T)

Dot-winged Antwren  Microrhopias quixensis
Occasionally seen at CC, usually near bamboo (T)

Striated Antbird  Drymophila devillei
Fairly common wherever there was extensive bamboo at CC, particularly around "Playa Bonita" (T)

Yellow-rumped Antwren  Terenura sharpei
A pair was seen on three occasions accompanying the canopy flock that roved around near the roadside just uphill of the Cock-of-the-Rock lek at SP; this species is classified as threatened, known from very few sites and with very few recent records in Peru (T)

Chestnut-shouldered Antwren  Terenura humeralis
Often heard in canopy flocks at CC, and probably present with all such flocks; only seen on c.10 occasions and difficult to see well (T)

Grey Antbird  Cercomacra cinerascens
Common in viny canopies at CC and PA (T)

Blackish Antbird  Cercomacra nigrescens
Only heard once in streamside tangles near "Playa Bonita", CC (T)

Black Antbird  Cercomacra serva
A few seen and heard in roadside shrubbery near SP (T)

Manu Antbird  Cercomacra manu
Fairly common in creekside bamboo at "Playa Bonita", CC (apparently not in the nearby upland bamboo), and in bamboo by the Madre de Dios river at PA (T)

White-browed Antbird  Myrmoborus leucophrys
Absent from floodplain forest at CC, but fairly common in terra firme and bamboo across the Manu river, and in the uplands near "Playa Bonita" (T)

Black-faced Antbird  Myrmoborus myotherinus
Common in floodplain forest and fairly common in terra firme at CC and PA (T)

Warbling Antbird  Hypocnemis cantator
Fairly common in terra firme and streamside tangles at CC and PA (T)

Band-tailed Antbird  Hypocnemoides maculicauda
Regularly found around the lakeshore at CC

Silvered Antbird  Sclateria naevia
Often seen and heard around the lakeshore at CC, and once along a stream through terra firme at PA (T)

White-lined Antbird  Percnostola lophotes
Fairly common, at least by voice, in bamboo near "Playa Bonita", CC and by the Madre de Dios river at PA (T)

Chestnut-tailed Antbird  Myrmeciza hemimelaena
Common at CC and PA (T)

Plumbeous Antbird  Myrmeciza hyperythra
Fairly common in creekside or swampy forest (especially where Heliconias abound) at CC (T)

Goeldi's Antbird  Myrmeciza goeldii
Common in swampy areas near the Manu and Madre de Dios rivers, especially in stands of Heliconia, at CC and PA (T)

Sooty Antbird  Myrmeciza fortis
Regular in drier floodplain and terra firme forest at CC and PA; often with antswarms (T)

Black-throated Antbird  Myrmeciza atrothorax
A few in scrappier habitat near the Manu and Madre de Dios rivers at CC (where common along the "Playa Bonita" and Ccolpa streams) and PA (T)

White-throated Antbird  Gymnopithys salvini
Small numbers regularly with antswarms at CC (T)

Hairy-crested Antbird  Rhegmatorhina melanosticta
1-3 regularly with antswarms at CC; two heard singing strongly along the Mirador trail at PA (T)

Spot-backed Antbird  Hylophylax naevia
Uncommon at CC, where encountered in undergrowth of floodplain forest on c.6 occasions (T)

Scale-backed Antbird  Hylophylax poecilinotus
Uncommon at CC, but seen and heard on a few occasions at PA (T)

Black-spotted Bare-eye  Phlegopsis nigromaculata
Regular with antswarms at CC (T)

Rufous-capped Antthrush  Formicarius colma
Regularly heard and occasionally seen at CC; once at an antswarm (T)

Black-faced Antthrush  Formicarius analis
Common by voice at CC and PA, and regularly seen (T)

Rufous-fronted Antthrush  Formicarius rufifrons
A few heard around

Rufous-breasted Antthrush  Formicarius rufipectus
Seen and heard around the Cock-of-the-Rock lek at SP, and along the loop trail next to the lodge (T)

Striated Antthrush  Chamaeza nobilis
Seen on 3 occasions at CC, and heard a few times more, but relatively scarce or non-vocal (T)

Undulated Antpitta  Grallaria squamigera
One heard singing at dusk and dawn near the hut at PI (T)

White-throated Antpitta  Grallaria albigula
One heard calling from dense vegetation on steep slopes around 1,700 m, above SP.

Red-and-white Antpitta  Grallaria erythroleuca
A few heard singing in roadside shrubbery near the huts at PI, where easily whistled into view

Stripe-headed Antpitta  Grallaria andicola
One seen well perched for a few minutes in lower branches of a Polylepis tree at AM

Amazonian Antpitta  Hylopezus berlepschi
Heard in tangled habitat near the stream at "Playa Bonita", CC

Thrush-like Antpitta  Myrmothera campanisona
Frequently heard and occasioally seen at dawn and dusk in floodplain and terra firme at CC and PA (T)

Ash-throated Gnateater  Conopophaga peruviana
Seen on c.6 occasions in the grid at CC

Rusty-belted Tapaculo  Liosceles thoracicus
Fairly common by voice in the grid at CC and also at PA; occasionally seen but only after playback (T)

TrillingTapaculo  Scytalopus parviristris
Heard only above PI

White-crowned Tapaculo  Scytalopus atratus
Heard regularly in roadside shrubbery above SP and seen once after playback (T)

Puna Tapaculo  Scytalopus simonsi
Seen and heard a few times in Polylepis at AM (T)

Cinereous Mourner  Laniocera hypopyrra
Fairly common and vocal at CC and PA in floodplain and terra firme (T)

Red-crested Cotinga  Ampelion rubrocristata
A few seen around PI

Barred Fruiteater  Pipreola arcuata
2-3 seen above PI

Screaming Piha  Lipaugus vociferans
Common at CC and PA (T)

Purple-throated Cotinga  Porphyrolaema porphyrolaema
A male seen briefly in the canopy, and a female seen well at close range accompanying a mid-stratum flock, both at CC

Plum-throated Cotinga  Cotinga maynana
Rarely seen in upper canopies at CC, chiefly because these are rarely visible, or flying over open areas

Spangled Cotinga  Cotinga cayana
One male seen flying over Cocha Totora, CC

Black-faced Cotinga  Conioptilon mcilhennyi
Fairly common and vocal (brrrrriiing) in floodplain forest at CC where two nests were found, one in the grid and one at "Playa Bonita" (T)

Bare-necked Fruitcrow  Gymnoderus foetidus
Fairly common at CC

Purple-throated Fruitcrow  Querula purpurata
Fairly common and vocal at CC (T)

Andean Cock-of-the-rock  Rupicola peruviana
Brilliant views of lekking males at SP, and regularly encountered in forest near there (T)

Band-tailed Manakin  Pipra fasciicauda
Several leks active in scrappier floodplain or swampy forest at CC (T)

Round-tailed Manakin  Pipra chloromeros
Scarce at CC and only seen at leks (usually in terra firme); more common at PA in similar habitat (T)

Blue-crowned Manakin  Lepidothrix coronata
Often seen and heard at CC and PA (T)

Yungas Manakin  Chiroxiphia boliviana
Fairly common and vocal in forest around SP (T)

Fiery-capped Manakin  Machaeropterus pyrocephalus
One male seen at CC, and the funny frog-like pink call heard occasionally

Dwarf Tyrant-Manakin  Tyranneutes stolzmanni
Common: males calling at many a song-post at CC and PA (T)

Wing-barred Piprites  Piprites chloris
Common at CC and PA, mostly with or near canopy flocks in terra firme (T)

Streak-necked Flycatcher  Mionectes striaticollis
Common in forest above SP

Olive-striped Flycatcher  Mionectes olivaceus
A few below SP along the Manu Road

Ochre-bellied Flycatcher  Mionectes oleagineus
Fairly common at CC and PA, where males were attending leks at both sites; we couldn't find Mionectes macconnelli (T)

Sepia-capped Flycatcher  Leptopogon amaurocephalus
A few seen around SP

Slaty-capped Flycatcher  Leptopogon superciliaris
Fairly common at CC and PA (T)

White-cheeked Tody-Tyrant  Poecilotriccus albifacies
Regularly found in bamboo at "Playa Bonita" and in uplands nearby, CC; also fairly common in bamboo near the Madre de Dios river at PA (T)

Flammulated Bamboo-Tyrant  Hemitriccus flammulatus
Fairly common in patches of bamboo in the uplands at CC, also in streamside cane and bamboo at "Playa Bonita" (T)

White-bellied Tody-Tyrant  Hemitriccus gresiepectus
Fairly common, at least by voice, in floodplain forest at CC, less common in terra firme hear and at PA (T)

Johannes' Tody-Tyrant  Hemitriccus iohannis
Only seen once: in tangled lower growth behind the lodge canteen at PA, where picked up by song (T)

Black-throated Tody-Tyrant  Hemitriccus granadensis
One seen and heard in stunted forest above PI (T)

Spotted Tody-Flycatcher  Todirostrum maculatum
Fairly common, at least by voice, in riverside habitat (Gynerium cane and Cecropia) at CC and just downriver from PA (T)

Yellow-browed Tody-Flycatcher  Todirostrum chrysocrotaphum
Regularly seen and heard in high canopies near the camp at CC; a few heard elsewhere and at PA but difficult to see (T)

Ringed Antpipit  Corythopis torquata
Seen or heard daily at CC and PA (T)

Bolivian Tyrannulet  Zimmerius bolivianus
Seemed fairly common at SP where picked up by plaintive peee call with canopy flocks, and sometimes in roadside shrubbery (T)

Slender-footed Tyrannulet  Zimmerius gracilipes
Seen rarely, and heard occasionally with canopy flocks at CC (T)

White-lored Tyrannulet  Ornithion inerme
Heard regularly at CC and occasionally at PA, mostly with canopy flocks (T)

Yellow-crowned Tyrannulet  Tyrannulus elatus
Common by voice at CC and PA (T)

Forest Elaenia  Myiopagis gaimardii
Fairly common by voice at CC and PA, and seen regularly with canopy flocks (T)

Yellow-bellied Elaenia  Elaenia flavogaster
A few seen around the lodge at SP (T)

Mottle-backed Elaenia  Elaenia gigas
2-3 in scrub along the river at PA

Highland Elaenia  Elaenia obscura
Two at MP

Sierran Elaenia  Elaenia pallatangae
Fairly common at PI

White-throated Tyrannulet  Mecocerculus leucophrys
A few in roadside flocks at PI

White-tailed Tyrannulet  Mecocerculus poecilocercus
Two in roadside flocks at PI

White-banded Tyrannulet  Mecocerculus stictopterus
Several in roadside flocks at PI and in stunted forest towards the treeline

Torrent Tyrannulet  Serpophaga cinerea
A few on rivers near SP

Tufted Tit-Tyrant  Anairetes parulus
Seen in shrubbery along the road between AM and OL

Ash-breasted Tit-tyrant  Anairetes alpinus
c.5 seen in Polylepis at AM

Marble-faced Bristle-Tyrant  Phylloscartes ophthalmicus
Fairly common and conspicuous component of mixed-species flocks near SP (T)

Cinnamon-faced Tyrannulet  Phylloscartes parkeri
A pair accompanying a mixed-species flock in tall trees in a scrubby clearing on the Manu Road below SP, c.1,000 m

Mottle-cheeked Tyrannulet  Phylloscartes ventralis
A few seen with mixed-species flocks in forest below the ruins at MP

Short-tailed Pygmy-Tyrant  Myiornis ecaudatus
Fairly common at CC; easily found once the voice is picked up (T)

Scale-crested Pygmy-Tyrant  Lophotriccus pileatus
Common in woodand and scrub along the Manu Road at SP (T)

Long-crested Pygmy-Tyrant  Lophotriccus eulophotes
Uncommon in bamboo in the uplands near "Playa Bonita," CC, where only seen on two occasions (using speculative playback) (T)

Large-headed Flatbill  Ramphotrigon megacephala
Fairly common in bamboo near "Playa Bonita", CC, and near the Madre de Dios river at PA (T)

Dusky-tailed Flatbill  Ramphotrigon fuscicauda
Not seen on the grid at CC, but encountered in tangled terra firme forest across the Manu river, and in bamboo in the uplands near "Playa Bonita". Also heard on several of occasions in bamboo on the Tinamou/Capybara trails, PA (T)

Rufous-tailed Flatbill  Ramphotrigon ruficauda
Often heard and regularly seen in tall forest at CC and PA (T)

Fulvous-breasted Flatbill  Rhynchocyclus fulvipectus
One seen in roadside shrubbery near Cloud Forest lodge, above SP

Zimmer's Flycatcher  Tolmomyias assimilis
Regularly heard with canopy flocks at CC, only seen well in the uplands near "Playa Bonita" (T)

Grey-crowned Flycatcher  Tolmomyias poliocephalus
Fairly common with canopy flocks at CC, especially near "Playa Bonita" (T)

Olive-faced Flycatcher  Tolmomyias viridiceps
Occasionally seen and heard with canopy flocks near the stream at "Playa Bonita", CC, and also in Cecropia dominated woodland near the Madre de Dios river at PA, and in clearing at c.1,000 m on the Manu Road below SP (T)

Golden-crowned Spadebill  Platyrinchus flavigularis
Regularly heard and occasionally seen in floodplain and terra firme at CC and PA (T)

White-crested Spadebill  Platyrinchus platyrhynchos
Encountered on c.10 occasions at CC, in both floodplain and terra firme (T)

Royal Flycatcher  Onychorhynchus coronatus
Never heard, and only seen twice with lower-stratum flocks at CC

Ornate Flycatcher Myiotriccus ornatus
c.3 seen on the Manu Road below SP, c.1,200 m (T)

Bran-colored Flycatcher  Myiophobus fasciatus
A few in streamside habitat at "Playa Bonita", CC (T)

Ruddy-tailed Flycatcher  Myiobius erythrurus
5-6 seen in floodplain forest at CC, and 1-2 seen along the Mirador trail at PA (T)

Cinnamon Flycatcher  Pyrrhomyias cinnamomea
Common along the Manu Road above SP (T)

Euler's Flycatcher  Lathrotriccus euleri
c.5 encountered at CC, but scarce (T)

Olive-sided Flycatcher  Contopus borealis
One seen perched on top of canopy of tall forest at PA

Smoke-colored Pewee  Contopus fumigatus
A few along the Manu Road near SP

Eastern Wood-Pewee  Contopus virens
1-2 along the road near SP identified by voice; other silent wood-pewees might have included sordidulus
Black Phoebe  Sayornis nigricans
Fairly common on rivers near SP

Vermilion Flycatcher  Pyrocephalus rubinus
Fairly common initially along rivers and lakeside at CC, but these birds left as the austral summer progressed

Rufous-breasted Chat-Tyrant  Ochthoeca rufipectoralis
Several along roadside at PI

Brown-backed Chat-Tyrant  Ochthoeca fumicolor
Several at the ruins above CU

White-browed Chat-Tyrant  Ochthoeca leucophrys
One at the ruins above CU

Drab Water-Tyrant  Ochthornis littoralis
Fairly common along the Manu and Madre de Dios rivers, as well as nearby streams (T)

Red-rumped Bush-Tyrant  Cnemarchus erythropygius
2-3 around the Polylepis groves at AM

Little Ground-Tyrant  Muscisaxicola fluviatilis
Regular on sand-bars along the Manu river

Plain-capped Ground-tyrant  Muscisaxicola alpina
Fairly common on the open valley floor at AM

Spot-billed Ground-tyrant  Muscisaxicola maculirostris
One at AM

White-winged Black-Tyrant  Knipolegus aterrimus
Several seen around PI and MP

Long-tailed Tyrant  Colonia colonus
A few in cleared areas on the Manu road below SP, at c.1,000 m

White-eyed Attila  Attila bolivianus
Common and vocal in floodplain forest, especially swampy areas, at CC (T)

Bright-rumped Attila  Attila spadiceus
Fairly common by voice at CC and PA, occurring in both floodplain and terra firme forest (T)

Greyish Mourner  Rhytipterna simplex
Fairly common from lower to upper-strata at CC and PA (T)

Sirystes  Sirystes sibilator
Fairly common and vocal with mixed-species flocks in the high canopy at CC (T)

Dusky-capped Flycatcher  Myiarchus tuberculifer
Fairly common on the Manu Road around SP (T)

Swainson's Flycatcher  Myiarchus swainsoni
Regularly seen with canopy flocks at CC (T)

Tropical Kingbird  Tyrannus melancholicus
Fairly commin in most habitats up to c.1,700 m, many nesting along rivers in lowlands (T)

Eastern Kingbird  Tyrannus tyrannus
Influxes of large numbers arrived along the Manu river in September

Crowned Slaty-Flycatcher  Griseotyrannus aurantioatrocristatus
Small numbers of migrants seen in high canopies along the "Playa Bonita" and Ccolpa streams, CC

Boat-billed Flycatcher  Megarynchus pitangua
Small numbers at CC and PA, usually along rivers or streams (T)

Lemon-browed Flycatcher  Conopias cinchoneti
Small groups encountered on c.3 occasions, foraging on top of canopies near the lodge at SP (T)

Golden-crowned Flycatcher  Myiodynastes chrysocephalus
Fairly common along the road above and below SP (T)

Streaked Flycatcher  Myiodynastes maculatus
Fairly common at CC and PA

Social Flycatcher  Myiozetetes similis
Common around lakes, rivers and streams in the lowlands, CC and PA (T)

Grey-capped Flycatcher  Myiozetetes granadensis
Fairly common in similar habitats to Social Flycatcher (T)

Piratic Flycatcher  Legatus leucophaius
Fairly common and vocal in canopies at CC and PA (T)

Lesser Kiskadee  Philohydor lictor
Uncommon around the lake at CC, commoner nearby at Cocha Totora (T)

Great Kiskadee  Pitangus sulphuratus
Fairly common around the lake at CC and in other edge habitats up to c.1,000 m on the Manu Road (T)

Várzea Schiffornis  Schiffornis major
Regularly heard and occasionally seen in swampy areas near the lake at CC (T)

Thrush-like Schiffornis  Schiffornis turdinus
Only heard at PA, on the Mirador trail

Barred Becard  Pachyramphus versicolor
1-2 seen accompanying mid-stratum flocks at SP

White-winged Becard  Pachyramphus polychopterus
Fairly common and vocal in riverine or streamside forest at CC and PA, rarely encountered in continuous tall forest (T)

Black-capped Becard  Pachyramphus marginatus
Uncommonly seen with canopy flocks at CC, but the distinctive dawn song regularly heard (T)

Pink-throated Becard  Pachyramphus minor
Fairly common at CC and PA, where regularly seen with canopy flocks

Black-tailed Tityra  Tityra cayana
Fairly common in swampy, riverine or streamside forest at CC

Masked Tityra  Tityra semifasciata
Regular in lakeside or riverside forest at CC and PA

Black-crowned Tityra  Tityra inquisitor
Occasionally seen at CC

White-collared Jay  Cyanolyca viridicyana
Fairly common around PI (T)

Purplish Jay  Cyanocorax cyanomelas
Fairly common in riverine forest along the Madre de Dios, near PA (T)

Violaceous Jay  Cyanocorax violaceus
Regularly heard and seen in tall floodplain forest at CC (T)

Green Jay  Cyanocorax yncas
Fairly common around and above SP (T)

Rufous-browed Peppershrike  Cyclarhis gujanensis
Fairly common along the Ccolpa and "Playa Bonita" streams at CC (Tt)

Slaty-capped Shrike-Vireo  Vireolanius leucotis
Commonly heard in canopy at CC and PA, but only seen once  (T)

Red-eyed Vireo  Vireo olivaceus
Fairly common with canopy flocks at CC  and PA, and in smaller numbers up to SP on the Manu Road (T)

Brown-capped Vireo  Vireo leucophrys
Fairly common with mixed-species flocks at SP (T)

Lemon-chested Greenlet  Hylophilus thoracicus
Never knowingly heard at CC, but a pair seen well with a mid-stratum flock at the uplands near "Playa Bonita"

Dusky-capped Greenlet  Hylophilus hypoxanthus
Common at CC and PA; heard with every canopy flock (T)

Tawny-crowned Greenlet  Hylophilus ochraceiceps
Commonly found with flocks of insectivores in lower-stratum at CC, usually picked up by its slow series of plaintive peeew notes (T)

White-capped Dipper  Cinclus leucocephalus
A pair seen on the river above SP

Andean Solitaire  Myadestes ralloides
Commonly heard and regularly seen along the road above SP (T)

White-eared Solitaire  Entomodestes leucotis
A few heard and c.5 seen above SP (c.1,800-2,000 m), including a pair with a nest (T)

Slaty-backed Nightingale-Thrush  Catharus fuscater
A few heard and one called in briefly with playback above SP (T)

Spotted Nightingale-Thrush  Catharus dryas
One heard around SP and another in song on the Mirador trail at PA (T)

Pale-eyed Thrush  Platycichla leucops
A few heard around SP, including around the Cock-of-the-Rock lek and 1-2 kms downhill of lodge. At this time of year seemed fairly easy to see as males were singing from open elevated perches (T)

Chiguanco Thrush  Turdus chiguanco
A few seen on the upper Manu Road and towards CU

Great Thrush  Turdus fuscater
Regularly seen on the upper Manu Road where common around PI (T)

Glossy-black Thrush  Turdus serranus
Fairly common on the upper Manu Road, especially around PI (T)

Black-billed Thrush  Turdus ignobilis
Common around BM

Lawrence's Thrush  Turdus lawrencii
Fairly common at CC and PA, usually in streamside and terra firme and less often heard in floodplain forest (T)

Hauxwell's Thrush  Turdus hauxwelli
Commonly seen and heard at CC and PA where it was the most vocal thrush; we definitely saw hauxwelli giving the distinctive "call" that is given under albicollis on the CDs of Voices of Amazonian Birds of South Peru and North Bolivia (T)

White-necked Thrush  Turdus albicollis
Regularly seen at CC and PA, but its more lackadaisical song only heard relatively infrequently (T)

Black-capped Donacobius  Donacobius atricapillus
Common around the lakes at CC and Totora (T)

Thrush-like Wren  Campylorhynchus turdinus
Small numbers around the lake at CC (T)

Grey-mantled Wren  Odontorchilus branickii
2-3 seen with canopy flocks just uphill of the Cock-of-the Rock lek at SP (T)

Moustached Wren  Thryothorus genibarbis
Common in streamside habitat and bamboo at CC and PA (T)

House Wren  Troglodytes aedon
Seen in riverside habitat along the Madre de Dios river at PA, also higher on Manu Road

Mountain Wren  Troglodytes solstitialis
A few seen with mixed-species flocks at PI

Grey-breasted Wood-Wren  Henicorhina leucophrys
Common and vocal in undergrowth along the Manu Road from well below SP, up to PI (T)

Southern Nightingale-Wren  Microcerculus marginatus
Fairly common in terra firme and floodplain forest in the grid at CC, and at PA, and once heard at c.1,300 m on the Manu Road (T)

Chestnut-breasted Wren  Cyphorhinus thoracicus
At least two pairs (groups?) heard and seen at SP, one on the loop trail by the lodge, another along the lower end of the Inca trail (which passes through very nice forest) (T)

Musician Wren  Cyphorhinus aradus
The amazing song was regularly heard and groups of 2-5 were quite frequently seen in the grid at CC (T)

Long-billed Gnatwren  Ramphocaenus melanurus
A few seen and heard in vine-tangled riverine habitat by the Manu river at CC (T)

Tropical Gnatcatcher  Polioptila plumbea
One pair heard singing and seen in canopies along the "Playa Bonita" stream at CC (T)

White-winged Swallow  Tachycineta albiventer
Common on lakes and rivers in the lowands, including CC and PA 

Brown-chested Martin  Phaeoprogne tapera
Fairly common on the lake at CC (where breeding in abandoned hornero nests around the lakeshore) and along major river in the lowlands (T)

Grey-breasted Martin  Progne chalybea
Regular along major rivers and occasionally seen over the lake at CC and nearby Totora; much commoner around settlements such as BM

Brown-bellied Swallow  Notiochelidon murina
A few seen at AM

Blue-and-white Swallow  Notiochelidon cyanoleuca
Flocks seen over the Madre de Dios river, PA, and up along the Manu Road to high altitudes

White-banded Swallow  Atticora fasciata
Fairly common along the Manu and Madre de Dios rivers, and on lakes and streams at CC

White-thighed Swallow  Neochelidon tibialis
A few seen along the "Playa Bonita" stream at CC

Southern Rough-winged Swallow  Stelgidopteryx ruficollis
Regularly seen along rivers and streams at CC and PA

Barn Swallow  Hirundo rustica
A few seen over the Madre de Dios river at PA

Tropical Parula  Parula pitiayumi
Fairly common in mixed-species flocks around SP (T)

Masked Yellowthroat  Geothlypis aequinoctialis
One pair seen at the "pantanal" near CC

Slate-throated Whitestart  Myioborus miniatus
Common in lower and mid-strata mixed-species flocks at SP, broadly overlapping with melanocephalus above the lodge (T)

Spectacled Whitestart  Myioborus melanocephalus
Fairly common in mixed-species flocks above SP, and occurring above the limit of miniatus towards and around PI (T)

Two-banded Warbler  Basileuterus bivittatus
Fairly common and vocal around SP, especially in bamboo patches downhill (T)

Pale-legged Warbler  Basileuterus signatus
Seen in stunted forest below the ruins at MP

Citrine Warbler  Basileuterus luteoviridis
Fairly common high on the Manu Road around PI

Russet-crowned Warbler  Basileuterus coronatus
Fairly common and vocal along the Manu Road below and above SP (T)

Three-striped Warbler  Basileuterus tristriatus
Common and conspicuous in lower-growth flocks along the Manu Road near SP (T)

Buff-rumped Warbler  Basileuterus fulvicauda
Common along rivers and especially streams at CC (T)

Rufous-collared Sparrow  Zonotrichia capensis
Common from PI to CU (T)

Yellow-browed Sparrow  Ammodramus aurifrons
Fairly common on grassy sand-banks along the Manu river

Pectoral Sparrow  Arremon taciturnus
Fairly common terra firme and bamboo at CC and PA (T)

Black-faced Brush-Finch  Atlapetes melanolaemus
Fairly common in lower-stratum roadside flocks above SP (T)

Tricolored Brush-Finch  Atalaptes tricolor
A few seen around PI

Stripe-headed Brush-Finch  Atlapetes torquatus
Two flew over the road at PI

Red-capped Cardinal  Paroaria gularis
Common around the lake at CC (T)

Bananaquit  Coereba flaveola
One pair in a mixed-species flock at c.1,000 m on the Manu road

Chestnut-vented Conebill  Conirostrum speciosum
One in a mixed-species flock at c.1,000 m on the Manu road

Cinereous Conebill  Conirostrum cinereum
Common between AM and OL

Giant Conebill  Oreomanes fraseri
One seen well in Polylepis at AM

White-browed Conebill  Conirostrum ferrugineiventre
One seen above OL

Red-billed Pied Tanager  Lamprospiza melanoleuca
Occasional in small groups accompanying mixed-flocks at CC

Magpie Tanager  Cissopis leveriana
Scarce at CC where seen mainly at "Playa Bonita". Also seen below SP

Grass-green Tanager  Chlorornis riefferii
A few at PI

Common Bush-Tanager  Chlorospingus ophthalmicus
Common between SP and PI (T)

Yellow-throated Bush-Tanager  Chlorospingus flavigularis
Common between SP and PI (T)

Black-capped Hemispingus  Hemispingus atropileus
A few at PI

Superciliaried Hemispingus  Hemispingus superciliaris
Regular in mixed flocks above SP and common at PI

Oleaginous Hemispingus  Hemispingus frontalis
One pair seen well in a mid-stratum flock at c.1,800 m, above SP

Black-eared Hemispingus  Hemispingus melanotis
One seen below PI

Orange-headed Tanager  Thlypopsis sordida
Seen once at the "pantanal" near CC

Rust-and-yellow Tanager  Thlypopsis ruficeps
Fairly common in roadside bushes around PI

Yellow-backed Tanager  Hemithraupis flavicollis
Small numbers regular in canopy flocks at CC and PA

Olive Tanager  Chlorothraupis carmioli
Fairly common along the mid and upper Mirador trail at PA (T)

White-winged Shrike-Tanager  Lanio versicolor
Common and vocal in canopy flocks at CC (T)

Slaty Tanager  Creurgops dentata
Seen a few times in small numbers above SP

Yellow-crested Tanager  Tachyphonus rufiventer
Fairly common in canopy flocks at CC and PA

Flame-crested Tanager Tachyphonus cristatus
Only seen once: a group of three in a canopy flock near Cocha Totora, CC

White-shouldered Tanager  Tachyphonus luctuosus
Common in mixed-species flocks at CC, PK, PA

Red-crowned Ant-Tanager  Habia rubica
Fairly common and vocal with lower-stratum flocks at CC and PA (T)

Scarlet Tanager  Piranga olivacea
Small numbers in mixed-species flocks below SP

Masked Crimson Tanager  Ramphocelus nigrogularis
Common around the lake at CC (T)

Silver-beaked Tanager  Ramphocelus carbo
Common around lakes and rivers in the lowlands (including CC and PA), up to SP (T)

Blue-grey Tanager  Thraupis episcopus
Small numbers at CC, BM, PA and below SP

Palm Tanager  Thraupis palmarum
Small numbers at PK, CC, BM, PA, and below SP

Blue-capped Tanager  Thraupis cyanocephala
Fairly common above SP and at PI

Blue-and-yellow Tanager  Thraupis bonariensis
Only seen (c.4 birds) at MP

Scarlet-bellied Mountain-Tanager  Anisognathus igniventris
Several seen well above SP and at PI (T)

Yellow-throated Tanager  Iridosornis analis
3-4 seen from 1,600-1,900 m, above SP

Golden-collared Tanager  Iridosornis jelskii
2-3 seen in low roadside bushes c.3 km above PI

Chestnut-bellied Mountain-Tanager  Delothraupis castaneoventris
A few seen from 1,700 m upwards between SP and PI

Fawn-breasted Tanager  Pipraeidea melanonota
Only seen once, above PI in elfin forest

Thick-billed Euphonia  Euphonia laniirostris
Regular at CC and common at PA (T)

White-lored Euphonia  Euphonia chrysopasta
Fairly common at CC where often heard, especially along the stream at "Playa Bonita" (T)

Bronze-green Euphonia  Euphonia mesochrysa
A few seen in lower or middle growth around SP

White-vented Euphonia  Euphonia minuta
Only seen twice in canopy flocks at CC

Orange-bellied Euphonia  Euphonia xanthogaster
Fairly common at CC, PA and SP (T)

Rufous-bellied Euphonia  Euphonia rufiventris
Common in canopy flocks at CC and PA (T)

Blue-naped Chlorophonia  Chlorophonia cyanea
A few in mixed-species flocks below SP

Orange-eared Tanager  Chlorochrysa calliparaea
Regular in mixed-species flocks above SP

Turquoise Tanager  Tangara mexicana
Fairly common around CC

Paradise Tanager  Tangara chilensis
Common at CC and PA, and below SP

Green-and-gold Tanager  Tangara schrankii
Very common at CC and PA, where several nests found

Golden Tanager  Tangara arthus
Common in canopy flocks above SP

Saffron-crowned Tanager  Tangara xanthocephala
Scarce in canopy flocks above SP

Golden-eared Tanager  Tangara chrysotis
Regular in mixed-species flocks above SP

Flame-faced Tanager  Tangara parzudakii
A few in mixed-species flocks above SP

Yellow-bellied Tanager  Tangara xanthogastra
Regular in small numbers in canopy flocks at CC and PA

Spotted Tanager  Tangara punctata
A few in mixed flocks at SP

Bay-headed Tanager  Tangara gyrola
Regular in small numbers in canopy flocks at CC and PA

Golden-naped Tanager  Tangara ruficervix
A few in mixed flocks above SP

Blue-necked Tanager  Tangara cyanicollis
A few around SP

Masked Tanager  Tangara nigrocincta
Fairly common in mixed flocks at CC

Beryl-spangled Tanager  Tangara nigroviridis
A few seen in mixed flocks above SP

Blue-and-black Tanager  Tangara vassorii
Very common above SP

Silver-backed Tanager  Tangara viridicollis
One male seen in a small mixed-flock below MP

Opal-rumped Tanager  Tangara velia
Small numbers seen in canopy flocks at CC, especially in the camp clearing

Opal-crowned Tanager  Tangara callophrys
Smal numbers regular in canopy flocks at CC

Black-faced Dacnis  Dacnis lineata
Regular at CC

Blue Dacnis  Dacnis cayana
Regular in canopy flocks at CC, PA and below SP

Tit-like Dacnis  Xenodacnis parina
2-3 in Polylepis at AM

Green Honeycreeper  Chlorophanes spiza
Regular in canopy flocks at CC, PA and below SP

Purple Honeycreeper  Cyanerpes caeruleus
Regular at CC and PA

Swallow-Tanager  Tersina viridis
Regular and breeding in sand-banks at the Ccolpa stream and "Playa Bonita" (T)

Peruvian Sierra-Finch  Phrygilus punensis
Small numbers seen at AM and CU

Ash-breasted Sierra-finch  Phrygilus plebejus
Fairly common in open areas around CU

Plumbeous Sierra-finch  Phrygilus unicolor
Common at AM

White-winged Diuca-finch  Diuca speculifera
c.4 near the pass at AM

Chestnut-breasted Mountain-finch  Poospiza caesar
One near the ruins at CU (in garden surrounded by tall pines), another flew across the road between OL and AM

Slate-colored Seedeater  Sporophila schistacea
Common and vocal in bamboo-dominated habitats towards the uplands on either side of the Manu river at CC, including at "Playa Bonita" (T)

Black-and-white Seedeater  Sporophila luctuosa
A few in bamboo and riverine cane at PK and CC

Yellow-bellied Seedeater  Sporophila nigricollis
A few at the ruins, MP

Chestnut-bellied Seedeater  Sporophila castaneiventris
Fairly common in open riverine shingle banks at PA

Black-billed Seed-Finch  Oryzoborus atrirostris
Only seen once: an adult in the swampy area at the near end of the lake at CC

Band-tailed Seedeater  Catamenia analis
A few at CU

Black-throated Flower-piercer  Diglossa brunneiventris
A few seen around CU

Deep-blue Flower-piercer  Diglossopis glauca
Fairly common above SP

Bluish Flower-piercer  Diglossopis caerulescens
A few seen and heard around 1,700 m, above SP (T)

Masked Flower-piercer  Diglossopis cyanea
Regular above SP (T)

Southern Yellow Grosbeak  Pheucticus chrysogaster
One pair seen at MP

Black-backed Grosbeak  Pheucticus aureoventris
One male seen at OL

Yellow-shouldered Grosbeak  Caryothraustes humeralis
Two were seen accompanying a mixed species flock in the uplands near "Playa Bonita", and another was seen with a mixed species flock in floodplain forest near the lake at CC

Slate-colored Grosbeak  Pitylus grossus
Rarely seen but often heard at CC (T)

Buff-throated Saltator  Saltator maximus
Fairly common in forest, particularly regrowing riverine areas, at CC (T)

Greyish Saltator  Saltator coerulescens
Fairly common at the lake at CC; also BM and along the river at PA (T)

Golden-billed Saltator  Saltator aurantiirostris
Only seen around OL

Blue-black Grosbeak  Cyanocompsa cyanoides
Often heard in undergrowth CC (especially along streams in terra firme) but rarely seen (T)

Casqued Oropendola  Psarocolius oseryi
Regularly seen along the stream at "Playa Bonita", CC, and also recorded at PA (T)

Crested Oropendola  Psarocolius decumanus
Uncommon at CC, apart from around "Playa Bonita" where fairly common. Also regular at PA and below SP in the bamboo zone (c.1,300 m) and lower (T)

Dusky-green Oropendola  Psarocolius atrovirens
At Cock-of-the-Rock lodge there is a Russet-backed Oropendola colony, whereas four km up the road the colony at the Cloud Forest lodge is this species. Common above SP (T)

Russet-backed Oropendola  Psarocolius angustifrons
Very common at CC, PA and around SP (T)

Olive Oropendola  Psarocolius yuracares
Apparently scarce at CC: only a few seen in the uplands area near "Playa Bonita"

Giant Cowbird  Molothrus oryzivora
Small numbers often seen flying over CC, and regular on sand-banks along major rivers

Yellow-rumped Cacique  Cacicus cela
Common throughout the lowlands (T)

Selva Cacique  Cacicus koepckeae
Small numbers regularly seen along the stream at "Playa Bonita", CC; generally picked up by their vocalisations: ching-ching chung-chung etc. (T)

Southern Mountain Cacique  Cacicus chrysonotus
Fairly common around PI (T)

Solitary Cacique  Cacicus solitarius
Fairly common but inconspicuous at CC. Often heard at "Playa Bonita" where birds were breeding (T)

Epaulet Oriole  Icterus cayanensis
Fairly common at CC and PA, usually associated with canopy flocks (T)

Troupial  Icterus icterus
Only encountered at "Playa Bonita" where this species was often heard along the stream and occasionally seen (T)

Pale-eyed Blackbird  Agelaius xanthophthalmus
Heard at the lake at CC but never seen there. It was easily seen nearby at the "pantanal" (T)

Velvet-fronted Grackle  Lampropsar tanagrinus
Seen on two occasions at the swampy area at the end of Cocha Cashu nearest the camp

Hooded Siskin  Carduelis magellanica
Regularly seen around CUZ

Olivaceous Siskin  Carduelis olivacea
Flocks regularly encountered along roadsides above SP

See some of Joes photos at www.worldbirder.com

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